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Swipe game, download game3574.ref061]\]. This difference in results reflects, in part, the choice of experimental techniques. Unlike manual tracking, which relies heavily on the definition of the region of interest, the CellProfiler application provides a comprehensive analysis of cell morphometrics, including quantification of morphology, size, and shape as well as per-cell analysis of color and texture.

In the interest of comparison with previous work, we have elected to include some of the quantitative results obtained by Davis et al. (2009). Specifically, in their study, they quantified 45–50 traits from a total of 8 single leukocytes and 2 separate leukocyte populations (tumor associated macrophages and T lymphocytes). One of the traits we measured (architectural asymmetry) is not shared with their study, while three of their traits (Circularity, Roundness, Feret min) were measured manually, while the rest of their traits were measured automatically. Despite the fact that our methods of measurement of these traits are not exact equivalents to theirs, the agreement between the two studies was quite high, with R-squared values of 0.88 for size traits, 0.76 for shape traits, and 0.94 for texture features.

Variations in Cell Morphology between Individuals {#sec031}
————————————————-

The variation in leukocyte morphology we observed is consistent with the known range of variation in neutrophil morphologies that exists between individuals \[[@pcbi.1003574.ref027], [@pcbi.1003574.ref062]\]. This is certainly not unexpected given that relatively little is known about the heritable component to neutrophil morphology. Moreover, although not entirely clear, the appearance of immature neutrophils is often a reliable predictor of age. However, given the well-known phenotypic difference in neutrophil morphology between healthy individuals of different age, it is perhaps surprising that we did not observe a significant correlation between age and cell shape.

Although we have attempted to control for several key potential confounders, such as disease state and date, our sample size is small. A larger dataset would be required to quantify the exact degree of phenotypic variation between healthy individuals, as well as to assess whether there are significant patterns in the demographics of the sample. Given the fact that neutrophils (and presumably other cell types) possess distinct subpopulations that are drawn from

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